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Carol McIntyre

Carol McIntyre


Carol McIntyre is a Wildlife Biologist with Denali National Park and Preserve, Alaska. She earned her B.S. from East Stroudsburg University, M.S. in Wildlife Management from University of Alaska-Fairbanks and Ph.D. in Wildlife Science from Oregon State University. She was elected a Fellow of the American Ornithologists' Union in 2016. Carol serves as an associate editor for the Journal of Raptor Research. chairs the Raptor Research Foundation (RRF) nominations committee, serves on the RRF Hamerstrom Award committee, and serves on the board of the Alaska Songbird Institute. When she isn’t in the field studying birds, she's out on the trail with her husband Ray and their small team of very large Alaska sled dogs.

National Park Service
4175 Geist Road  
Fairbanks,  Alaska  99709  United States
Phone: 907-455-0671
Email: Carol_McIntyre@nps.gov

   Research Interests

I am interested in the response of northern breeding raptors to changes and events on their breeding grounds, migration corridors, and wintering areas, including the cascading effects of climate change.

Species of Interest: 
  • Osprey
  • Bald Eagle
  • Northern Harrier
  • Red-tailed Hawk
  • Rough-legged Hawk
  • Golden Eagle
  • American Kestrel
  • Merlin
  • Gyrfalcon
  • Peregrine Falcon
  • Great Horned Owl
  • Snowy Owl
  • Northern Hawk Owl
  • Great Gray Owl
  • Short-eared Owl
  • Boreal Owl
  • Northern Saw-whet Owl


Mizel, J.D., J. H. Schmidt, and C. L. McIntyre. 2021. Climate and weather have differential effects in a high-latitude passerine community. Oecologia.

Davidson, S. C., G. Bohrer, E. Gurarie, S. LaPoint, P. J. Mahoney, ... K. Joly, ... J. P. Lawler, ...B. Mangipane, ... C. L. McIntyre, ... P. A. Owen, ... M. S. Sorum, ... et al. 2020. Ecological insights from three decades of animal movement tracking across a changing Arctic. Science 370(6517): 712-715.

Franke, A., K. Hawkshaw, S. Ambrose, ... C. L. McIntyre, et al. 2020. Status and trends of circumpolar peregrine falcon and gyrfalcon populations. Ambio 49(3): 762-783.

Stenhouse, I. J., E. M. Adams, L. M. Phillips, S. Weidensaul, and C. L. McIntyre. 2019. A preliminary assessment of mercury in the feathers of migratory songbirds breeding in the North American subarctic. Ecotoxicology, DOI 10.1007/s10646-019-02105-2.

Mizel, J. D., Schmidt, J. H., Phillips, L. M. and Mcintyre, C.L., 2019. A binomial N‐mixture model for estimating arrival and departure timing. Methods in Ecology and Evolution 10(7): 1062-1071.

DeLuca, W. V., B. K. Woodworth, S. A. Mackenzie, A. E. M. Newman, H. A. Cooke, L. M. Phillips, N. E. Freeman, A. O. Sutton, L. Tauzer, C. McIntyre, I. J. Stenhouse, S. Weidensaul, P. D. Taylor, and D. R. Norris. 2019. A boreal songbird’s 20,000 km migration across North America and the Atlantic Ocean. Ecology 00(00):e02651. 10.1002/ecy.2651

Eisaguirre, J. M., T. L. Booms, C. P. Barger, C. L. McIntyre, S. B. Lewis, and G. A. Breed. 2018. Local meteorological conditions reroute a migration. Proceedings Royal Society B 285:20181779.

Mizel, J. D.,C.L. McIntyre, S.B. Lewis, M.S. Lindberg, and J.H. Schmidt. 2018. A multi‐state, time‐removal model for population dynamics of cliff‐nesting raptors. Journal of Wildlife Management 82:1701-1710. doi:10.1002/jwmg.21526

Fuglei, E., J. A. Henden, C. T. Callahan, O. Gilg, J. Hansen, R. A. Ims, A. P. Isaev, J. Lang, C. L. McIntyre, R. A. Merizon, O. Y. Mineev, Y. N. Mineev, D. Mossop, O. K. Nielsen, E. B. Nilsen, A. O. Pedersen, N. M. Schmidt, B. Sittler, M. H. Willebrand, and K. Martin. 2018. Circumpolar status of Arctic ptarmigan: Population dynamics and trends. Ambio DOI 10.1007/s13280-019-01191-0.

Mizel, J. D., J. H. Schmidt, C. L. McIntyre, and M. S. Lindberg. 2017. Subarctic-breeding passerines exhibit phenological resilience to extreme spring conditions. Ecosphere 8(2):e01680. 10.1002/ecs2.1680

Schmidt, J.H., E.A. Rexstad, C.A. Roland, C.L. McIntyre, M.C. MacCluskie, and M.J. Flamme. 2017. Weather-driven change in primary productivity explains variation in the amplitude of two herbivore population cycles in a boreal system. Oecologia DOI 10.1007/s00442-017-4004-3

Schmidt, J.H., C.L. McIntyre, C.A. Roland, M.C. MacCluskie, and M.J. Flamme. 2017. Bottom-up processes drive reproductive success of an apex predatory. Ecology and Evolution. DOI: 10.1002/ece3.3800

Steenhof, K., M.N. Kochert, C.L. McIntyre and J.L. Brown. 2017. Coming to terms about describing golden eagle reproduction. Journal of Raptor Research 51: 378-390.

Brown, J.L., B. Bedrosian, D.A. Bell, M.A. Braham, J. Cooper, R.H. Crandall, J. DiDonato, R. Domenech, A.E Duerr, T. E. Katzner, M. J. Lanzone, D.W. LaPlante, C.L. McIntyre, T.A. Miller, R.K. Murphy, A. Shreading, S.J. Slater, J.P. Smith, B.W. Smith, J.W. Watson and B.Woodbridge. 2017. Patterns of spatial distribution of golden eagles across North America: how do they fit into existing landscape-scale mapping patterns? Journal of Raptor Research 51: 197-215.

Franke, A., C.L. McIntyre, and K. Steenhof. Terminology. In Applied Raptor Ecology: essentials from gyrfalcon research. Edited by D. Anderson, C. McClure, and A. Franke.

McIntyre, C.L. and S.B. Lewis. 2016. Observations of migrating Golden Eagles in eastern interior Alaska offer insights on population size and migration monitoring. Journal of Raptor Research 50: 254-264.

Mizel, J. D., J. H. Schmidt, C. L. Mcintyre, and C. A. Roland. 2016. Rapidly shifting elevational distributions of passerine species parallel vegetation change in the subarctic. Ecosphere 7(3):e01264. 10.1002/ecs2.1264

Doyle, K.M., T.E. Katzner, G.W. Roemer, J.W. Cain, III, B.A. Millsap, C.L. McIntyre, S.A. Sonsthagen, N.B. Fernandez, M. Wheeler, Z. Bulut, P.H. Bloom and J.A. DeWoody. 2016. Genetic structure and viability selection in the golden eagle (Aquila chrysaetos), a vagile raptor with a Holarctic distribution. Conservation Genetics. July 2016.

McIntyre, C.L., and M.D. Paulson. 2015. What came first, the nest or the egg? An unusual golden eagle nest observed in Denali National Park and Preserve, Alaska. Journal of Raptor Research 49: 98-101.

Fackler, P.L., K. Pacific, J. Martin, and C. McIntyre. 2014. Efficient use of information in adaptive management with an application to managing recreation near golden eagle nesting sites. PLoS ONE 9(8): e102434. doi:10.1371/journal.pone.0102434.

Schmidt, J.H., C.L. McIntyre, and M.C. MacCluskie. 2013. Accounting for incomplete detection: what are we estimating and how might it affect long-term passerine monitoring programs. Biological Conservation 160: 130-139.

McIntyre, C.L., and J.H. Schmidt. 2012. Ecological and environmental correlates of territory occupancy and breeding performance of migratory Golden Eagles Aquila chrysaetos in interior Alaska. Ibis 154:124-135.

Martin, J., P.L. Fackler, J.D. Nichols, M.C. Runge, C.L. McIntyre, B.L. Lubow, M.C. McCluskie, and J.A. Schmutz. 2011. An adaptive-management framework for optimal control of hiking near Golden Eagle nests in Denali National Park. Conservation Biology 25:316-323.

Martin, J., C.L. McIntyre, J.E. Hines, J.D. Nichols, J.A. Schmutz, and M.C. MacCluskie. 2009. Dynamic multistate site occupancy models to evaluate hypotheses relevant to conservation of Golden Eagles in Denali National Park, Alaska. Biological Conservation 142:2726-2731.

McIntyre, C.L., J.M. Withers, and N.J. Gates. 2009. Merlins (Falco columbarius) scavenge road-killed Snowshoe Hares (Lepus americanus) in interior Alaska. Journal of Raptor Research 43:254.       pdf

McIntyre, C.L., D.C. Douglas, and L.G. Adams. 2009. Movements of juvenile Gyrfalcons from western and interior Alaska following departure from their natal areas. Journal of Raptor Research 43:99-109.       pdf

McIntyre, C.L., D.C. Douglas, and M.W. Collopy. 2008. Movements of Golden Eagles (Aquila chrysaetos) from interior Alaska during their first year of independence. Auk 125:214-224.        pdf

McIntyre, C.L., and M.W. Collopy. 2006. Postfledging dependence period of migratory Golden Eagles (Aquila chrysaetos) in Denali National Park and Preserve, Alaska. Auk 123:877-884.

McIntyre, C.L., M.W. Collopy, J.G. Kidd, and A.A. Stickney. 2006. Characteristics of the landscape surrounding Golden Eagle nest sites in Denali National Park and Preserve, Alaska. Journal of Raptor Research 40:46-51.       pdf

McIntyre, C.L., M.W. Collopy, and M.S. Lindberg. 2006. Survival probability and mortality of migratory juvenile Golden Eagles from Interior Alaska. Journal of Wildlife Management 70:717-722.

McIntyre, C., K. Steenhof, M.N. Kochert and M.W. Collopy. 2006. Long-term Golden Eagle studies in Denali National Park and Preserve. Alaska Park Science 5:42-45.

Brown, R.J., and C. McIntyre. 2005. New prey species documented for Northern Pike (Esox lucius): Bald Eagle (Haliaeetus leucocephalus). Arctic 58:437.

McCaffery, B.J., and C. McIntyre. 2005. Disparities between results and conclusion: do Golden Eagles warrant special concern based on migration counts in the western United States? Condor 107:469-473.

McIntyre, C.L. 2002. Patterns in nesting area occupancy and reproductive success of Golden Eagles (Aquila chrysaetos) in Denali National Park and Preserve, Alaska, 1988-99. Journal of Raptor Research 36:50-54.

McIntyre, C.L., and L.G. Adams. 1999. Reproductive characteristics of migratory Golden Eagles in Denali National Park, Alaska. Condor 101:115-123.

McIntyre, C.L., and R.E. Ambrose. 1999. Raptor migration in autumn through the Upper Tanana River Valley, Alaska. Western Birds 30:33-38.

Young, D.D., Jr., C.L. McIntyre, P.J. Bente, T.R. McCabe, and R.E. Ambrose. 1995. Nesting by Golden Eagles on the north slope of the Brooks Range in northeastern Alaska. Journal of Field Ornithology 66:373-379.

Swem, T., C. McIntyre, R.J. Ritchie, P.J. Bente, and D.G. Roseneau. 1994. Distribution, abundance, and notes on the breeding biology of Gyrfalcons Falco rusticolus in Alaska. Pp. 437-444 in B.-U. Meyburg and R.D. Chancellor (eds.), Raptor conservation today. World Working Group on Birds of Prey, Berlin, and Pica Press, London.

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